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How dreadfully sad for you. Sometimes there simply are no answers, and no matter what you do or how hard you try, the outcome can not be changed. There can be problems that date back generations, but they simply lie dormant , then without any foreseeable reason, they suddenly rear up. You clearly loved and cared for your little ones, and gave them lives that many, many animals can only ever dream of
Thank you. I've accepted now I probably won't ever have an answer, which is hard but it is what it is.
My other gerbils Astra and Twiglet are still well, though very upset that I'm not giving them the attention they're used to. Most germs don't live more than 2 weeks in the environment so I'm going to give it that long before handling them again. My hamster Kebab is also well. He's very old now so doesn't often want handling anyway.